The world's oldest known gorilla has died peacefully in her sleep, just a month after celebrating her 60th birthday.
At her death, Colo, the first gorilla to be born in captivity, had outlived the average gorilla by 20 to 30 years, according to her keepers.
In her long time on Earth, Colo became the matriarch of a huge gorilla family, having lived to see the birth of three children, 16 grandchildren, 12 great-grandchildren and three great-great-grandchildren, according to the Columbus Zoo and Aquarium, where she lived.
On Dec. 19, thousands of people, both at the zoo and online, celebrated the beginning of her seventh decade with a big party, complete with a decorated enclosure, special presents and a custom, gorilla-friendly cake.
"Colo touched the hearts of generations of people who came to see her and those that cared for her over her long lifetime," Tom Stalf, president and CEO of the Columbus Zoo and Aquarium, said in a statement. "She was an ambassador for gorillas and inspired people to learn more about the critically endangered species and motivated them to protect gorillas in their native habitat." [Photos: Colo, World's Oldest Gorilla, Turns 60]
Colo was born at the Columbus Zoo on Dec. 22, 1956, at a time when little was known about keeping and raising western lowland gorillas in captivity, according to a statement from the zoo. She was born a preemie, and a zookeeper found her still in her amniotic sac in a zoo enclosure, after having been abandoned by her mother, according to a 2009 video about her by Columbus Zoo Media. Thanks to 24/7 attention by her human caregivers, she survived and thrived. Colo was briefly called "Cuddles," but after a naming contest, was given her current moniker — a combination of "Columbus" and "Ohio."
In 1958, she met a male gorilla named Bongo, who became her mate. The duo's "love affair" outlasted many human marriages, continuing for 25 years.
Though Colo spent her life near other gorillas, as she aged, she seemed to do better with more personal space, and so she was moved to a private enclosure. She was also plagued by arthritis in her later years, Audra Meinelt, the assistant curator of the Congo Expedition at the Columbus Zoo, previously told Live Science.
In addition to dietary supplements to prevent joint stiffness, Colo was also given a kind of gorilla physical therapy to encourage her to stay active.
"Her arthritis is very specific to her hands and feet, so we came up with ways to get her to use her fingers more often," Meinelt said. "We also changed the way that we present her diet, so that also causes her to exercise her fingers," she said.
Colo also became choosier about her food, shunning green beans, grapes, zucchini, honeydew, oranges, cantaloupe and strawberries as she got older, Dana Hatcher, manager of animal nutrition at the Columbus Zoo, previously told Live Science. To make sure that she got ample nutrition, her caregivers also gave Colo probiotics, brewer's yeast for B vitamins and iceberg lettuce for its water content.
Originally published on Live Science.
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Tia is the managing editor and was previously a senior writer for Live Science. Her work has appeared in Scientific American, Wired.com and other outlets. She holds a master's degree in bioengineering from the University of Washington, a graduate certificate in science writing from UC Santa Cruz and a bachelor's degree in mechanical engineering from the University of Texas at Austin. Tia was part of a team at the Milwaukee Journal Sentinel that published the Empty Cradles series on preterm births, which won multiple awards, including the 2012 Casey Medal for Meritorious Journalism.