Fossil Record: A Gallery of "Bugged" Medieval Books

A woodblock dotted with holes made by furniture beetles. Marred woodblocks left behind white dots on the printed page, providing a record of European beetle ranges, according to research published Nov. 20 in the journal Biology Letters.
The 1541 woodcut "De Rijke Man" (The Rich Man) by Cornelius Anthonisz. White circles reveal where furniture beetles gnawed through the wood block before printing.
By tracking wormholes such as the ones that appear on this print of "The Rich Man," Pennsylvania Statue University biologist Blair Hedges was able to determine where different species lived at the time.
Anobium punctatum, the common furniture beetle, is responsible for wormholes in woodcuts from northern Europe.
This diagram shows how furniture beetles bore through dry wood. Larvae hatch and crawl out of the wood in their adult forms.
This map shows the medieval distribution of the common furniture beetle (northern Europe, in blue) and the Mediterranean furniture beetle (southern Europe, in red). Today, the beetles coexist across western Europe.
This woodcut depicts a carver creating a woodblock for use in printing. Most first-edition books show little evidence of beetle damage, but later editions often show holes as beetles move in and infest the stored blocks.
In Asia, the art of woodcut printing began even earlier than in Europe. This colorful print of Buhddha was made in the 10th Century in Shanxi, China.
